Lily Owens - The 14-year-old white protagonist and narrator. She is motherless and escapes her abusive father T. Ray with her nanny Rosaleen, seeking refuge with the Boatwright sisters and answers about her late mother.

Rosaleen Daise - Lily's fierce-hearted, proud, and outspoken black nanny who is a surrogate mother to Lily. She is arrested for insulting racists, prompting their escape.

August Boatwright - The warm, maternal leader of the three beekeeping sisters who take in Lily and Rosaleen. She helps Lily come to terms with her mother's past.

June Boatwright - August's protective and initially hostile younger sister who eventually warms up to Lily.

May Boatwright - The youngest, trauma-haunted Boatwright sister with a special connection to the Black Madonna.

Zach Taylor - A young black man who works on the Boatwright farm and becomes Lily's friend and love interest.

T. Ray Owens - Lily's cold, abusive father who lies about the circumstances of Lily's mother Deborah's death.

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