Ella Turner: A modern-day American woman who moves to the small French village of Lisle-sur-Tarn with her husband Rick. Ella struggles to fit in and begins experiencing strange dreams and events that connect her to her 16th-century ancestor, Isabelle.

Isabelle du Moulin: Also known as "La Rousse" due to her red hair. Isabelle is a young peasant girl in 16th-century France. She marries Etienne Tournier, who comes from a Huguenot family. Isabelle's story is told in alternating chapters alongside Ella's.

Etienne Tournier: Isabelle's husband, a member of a Huguenot family in 16th-century France. When the Tourniers face threats from Catholic neighbors, they flee to Geneva.

Jean-Paul: The charming librarian in Lisle-sur-Tarn who befriends Ella and helps her research her family history.